Jiangtao Zhou a, b, c, d, 1
These authors contributed equally to this work.
, Yiqun Chen a, b, c, d, 1
These authors contributed equally to this work.
, Zheng Shao a, b, c, d, Shiping Ding a, b, c, d, Wenbao Qi a, b, c, d, Jiahao Zhang a, b, c, d, ⁎
, Lihong Huang a, b, c, d, ⁎ 
a Guangdong Laboratory for Lingnan Modern Agriculture, College of Veterinary Medicine, South China Agricultural University, Guangzhou 510642, China
b National Avian Influenza Para-Reference Laboratory, Guangzhou 510642, China
c National Development and Reform Commission of People's Republic of China, National and Regional Joint Engineering Laboratory for Medicament of Zoonoses Prevention and Control, Guangzhou 510642, China
d Key Laboratory of Zoonoses, Ministry of Agriculture and Rural Affairs of the People's Republic of China, Guangzhou 510642, China