Jinyue Guo a, 1
These author contributed equally to this work.
, Hai Yu b, 1
These author contributed equally to this work.
, Congying Wang a, 1
These author contributed equally to this work.
, Sheng Yuan a, Hailiang Sun c, Jing Yang a, Qing Cheng a, Kaijian Luo c, Yong Li d, Zhili Li a, Quan Liu a, Shujian Huang a, Feng Wen a, ⁎ 
a College of Life Science and Engineering, Foshan University, No33 Guangyun road, Shishan town, Nanhai District, Foshan, Guangdong 528231, China
b Shanghai Veterinary Research Institute, Chinese Academy of Agricultural Sciences, Shanghai 200241, China
c College of Veterinary Medicine, South China Agricultural University, Guangzhou, Guangdong 510642, China
d College of Animal Science and Technology, Jiangxi Agricultural University, Nanchang, Jiangxi 330045, China