Jiahao Zhang a, b, e, f, 1
These authors contributed equally to this work.
, Lihong Huang d, 1
These authors contributed equally to this work.
, Yiqun Chen a, b, e, f, Xiaomin Wang a, b, e, f, Ming Liao a, b, c, e, f, g, h, ⁎
, Wenbao Qi a, b, c, e, f, g, h, ⁎ 
a College of Veterinary Medicine, South China Agricultural University, Guangzhou 510642, PR China
b National Avian Influenza Para-reference Laboratory, Guangzhou 510642, PR China
c Guangdong Laboratory for Lingnan Modern Agriculture, Guangzhou 510642, PR China
d Department of Biomedical Sciences, City University of Hong Kong, Hong Kong 999077, PR China
e National and Regional Joint Engineering Laboratory for Medicament of Zoonoses Prevention and Control, National Development and Reform Commission of the People's Republic of China, Guangzhou 510642, PR China
f Key Laboratory of Zoonoses, Ministry of Agricultural and Rural Affairs of the People's Republic of China, Guangzhou 510642, PR China
g Key Laboratory of Animal Vaccine Development, Ministry of Agricultural and Rural Affairs of the People's Republic of China, Guangzhou 510642, PR China
h Key Laboratory of Zoonoses Prevention and Control of Guangdong Province, Guangzhou 510642, PR China