Jiahao Zhang 1
These authors contributed equally to this work.
, Guangjie Lao 1
These authors contributed equally to this work.
CNational and Regional Joint Engineering Laboratory for Medicament of Zoonoses Prevention and Control, College of Veterinary Medicine, South China Agricultural University, Guangzhou, Guangdong 510642, PR China
Ronghua Zhang
Center for Animal Disease Control and Prevention, Honghe, Yunnan 654400, PR China
Zhengji Wei
Center for Animal Disease Control and Prevention, Liuzhou, Guangxi 545005, PR China
Hexing Wang
Center for Animal Disease Control and Prevention, Honghe, Yunnan 654400, PR China
Guanming Su
National and Regional Joint Engineering Laboratory for Medicament of Zoonoses Prevention and Control, College of Veterinary Medicine, South China Agricultural University, Guangzhou, Guangdong 510642, PR China
Nan Shan
Nanjing Institution of Environmental Sciences, Ministry of Environmental Protection of China, Nanjing, Jiangsu 210093, PR China
Bo Li, Huanan Li
National and Regional Joint Engineering Laboratory for Medicament of Zoonoses Prevention and Control, College of Veterinary Medicine, South China Agricultural University, Guangzhou, Guangdong 510642, PR China
Yuandi Yu
Chongqing Academy of Animal Sciences, Chongqing 402460, PR China
Weixin Jia, Ming Liao *
, Wenbao Qi * 
National and Regional Joint Engineering Laboratory for Medicament of Zoonoses Prevention and Control, College of Veterinary Medicine, South China Agricultural University, Guangzhou, Guangdong 510642, PR China
Key Laboratory of Zoonoses, Key Laboratory of Animal Vaccine Development, Ministry of Agriculture, Guangzhou, Guangdong 510642, PR China
Key Laboratory of Zoonoses Prevention and Control of Guangdong Province, Guangzhou, Guangdong 510642, PR China